Caupedactylus, Lower Cretaceous. Caupedactylus was found in limestone that formed under the waves of the incipient Atlantic Ocean, but lived close enough to the shore that it was probably not fully marine in its habits. Tapejarids are thought to have been omnivorous, eating fish, invertebrates, and small terrestrial vertebrates, as well as fruit, seeds, and other plant material. It's likely that Caupedactylus was similar in its habits, and would have had a wide variety of food at its disposal.